Marsilea vestita
hairy clover-fern, pepperwort, water-clover pepperwort

Distribution: Occurring chiefly east of the Cascades crest and in the Columbia River Gorge in Washington; British Columbia to California, east to the Great Plains and southeastern U.S.

Habitat: Ponds, vernal pools, and floodplains from low to middle elevations.

Spores: April-October

Origin: Native

Growth Duration: Perennial

Conservation Status: Not of concern


Rhizomatous perennials forming clones, aquatic to amphibious; stems creeping on substrate surface or subterranean; roots emerging from nodes.


Leaves long-petiolate, petiole threadlike, 2-20 cm, slightly pubescent; blade palmately divided into 4 pinnae, pinnae obdeltate, 4-19 mm long and 4-16 mm broad, pubescent to glabrous.


Sporocarp stalks erect and not branched, attached at base of petiole or barely above it, not hooked at apex, 0.5-25 mm; sporocarps at right angle to stalk or slightly nodding, 3.6-7.6 mm long and 1.5-2 mm broad, elliptic to nearly round in lateral view, pubescent but becoming glabrous, scars from dropped trichomes commonly appearing as small purple or brown specks; sori 14-22.

Accepted Name:
Marsilea vestita Hook. & Grev.
Publication: Icon. Filic. 2: plate 159. 1830.

Synonyms & Misapplications:
Marsilea fournieri C. Chr.
Marsilea mucronata A. Braun
Marsilea tenuifolia Engelm. ex A. Braun
Marsilea uncinata A. Braun
Marsilea vestita Hook. & Grev. ssp. tenuifolia (Engelm. ex A. Braun) D.M. Johnson
Marsilea vestita Hook. & Grev. ssp. vestita [JPM]
Additional Resources:

PNW Herbaria: Specimen records of Marsilea vestita in the Consortium of Pacific Northwest Herbaria database

WA Flora Checklist: Marsilea vestita checklist entry

OregonFlora: Marsilea vestita information

E-Flora BC: Marsilea vestita atlas page

CalPhotos: Marsilea vestita photos

34 photographs:
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